Checklist of the caddisflies (Insecta, Trichoptera) of the Upper Midwest region of the United States

David C. Houghton, R. Edward Dewalt, Todd Hubbard, Kurt L. Schmude, Jeffrey J. Dimick, Ralph W. Holzenthal, Roger J Blahnik, James L. Snitgen

Research output: Contribution to journalArticlepeer-review

6 Scopus citations


Five hundred and fifty-two caddisfly species are reported from the Upper Midwest region of the United States, an area that includes 13 states and ~ 2 million km2. Of these, 62 species are reported for the first time from the state of Iowa, 25 from Wisconsin, 18 from South Dakota, 12 from Illinois, five from Indiana, four from North Dakota, four from Minnesota, and one from Nebraska. The Upper Midwest fauna contains nearly 40% of all species known from the United States and Canada, as well as 22 species endemic to the region. Overall species richness was highest in Michigan (319 species), Kentucky (296), Minnesota (292), and Wisconsin (284). Differences in state species assemblages within the region largely followed a geographic pattern, with species richness declining in the western prairie states. There are almost certainly further species remaining to be found in this large region.

Original languageEnglish (US)
Pages (from-to)287-300
Number of pages14
Issue number1111
StatePublished - 2022

Bibliographical note

Funding Information:
The sampling of Sleeping Bear Dunes National Lakeshore was conducted under permit SLBE-2014-SCI-0002, facilitated by Kevin Skerl. The Saint Croix National Scenic Waterway was sampled under permits SACN-2013-SCI-0003 and SACN-2016-SCI-0001, facilitated by Jill Midland. The staff at Pictured Rocks National Lakeshore granted access to streams under permits PIRO-2010-0008 and PIRO-2013-0002. Permission to sample in the Huron Mountains of Michigan was provided by the Huron Mountain Wildlife Foundation. Permission to sample at the Black River Ranch of Michigan was provided by the Black River Ranch Board of Directors. Permission to sample at Sarah Jane’s Natural Area of Michigan was provided by John Bagley and Andrew Bacon (Michigan Nature Association). Google Earth base maps were used following permission guidelines (https://www. The valuable comments of Desiree Robertson and Paul Frandsen improved earlier version of the manuscript. This is paper #30 of the G.H. Gordon BioStation Research Series.

Funding Information:
Primary funding for this research came from a U.S. Environmental Protection Agency Science to Achieve Results Fellowship and Minnesota Natural Heritage and Nongame Research Program grant to DCH, and from U.S. Department of Interior (INT RD X-1-R-1), National Science Foundation (DEB 09-18805 ARRA), Illinois State Wildlife (IDNR FWS T-121-R-1), and Indiana Department of Natural Resources (E16-21-40777, 0017556043) grants to RED. Further support came from the Hillsdale College (HC) biology department, grants from the Huron Mountain Wildlife Foundation, several HC LAUREATES grants to DCH and affiliated students, and a HC Faculty Summer Leave grant. Funding to support KLS for surveys of state-owned properties in Wisconsin came from the Wisconsin Department of Natural Resources, Natural Heritage Inventory.

Publisher Copyright:
© 2022 David C. Houghton et al.


  • Caddisfly
  • Midwest
  • Trichoptera
  • USA
  • checklist
  • diversity


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